- Investigator, Howard Hughes Medical Institute
- Chairman and Professor of Genome Sciences and Professor of Medicine, University of Washington
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- Professor of Anesthesia and Pharmaceutical Chemistry, UCSF
- Member, Institute of Medicine of the National Academy of Sciences
- Member, UCSF Comprehensive Cancer Center
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- Professor, Banting and Best Department of Medical Research, University of Toronto
- Associate Investigator, Ontario Institute for Cancer Research
- Led development of phage display technology at Genentech, Inc.

- Professor of Biological Engineering and Computer Science, Massachusetts Institute of Technology
- Co-founder and co-chair, Computational and Systems Biology Initiative at Massachusetts Institute of   Technology
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